The story begins with the protagonist, a male "Hero" who has endured over a century of failed attempts to defeat the Demon King. Each death resets his life to age fifteen, but he retains all memories and the crushing weight of past failures and betrayals. His latest life ends in a desperate battle for the capital, where he is assassinated by a Royal Guard knight, confirming his cynical belief that the kingdom would betray him.
However, this time, his reincarnation is different. He awakens in a forest as a young woman, stripped of his Heroic powers. A sarcastic envoy named Jane, speaking from a Goddess statue, explains that a "Rebellious God" interfered with his reincarnation, forcing his soul into a new, weaker female body in a parallel world. No longer the chosen Hero, and with humanity facing extinction in five years, the protagonist, now calling herself Meme (a name symbolizing his first major sin), remains consumed by the singular goal of killing the Demon King, now a personal vendetta rather than a divine duty.
Struggling with her physically weaker body and new identity, Meme encounters her past self, Oscar, who is now the new, naive fifteen-year-old Hero, wielding the Holy Sword. Accompanying him is Karen, his kind-hearted childhood friend and a talented Holy Magic user. Meme, despite her cynicism and the emotional turmoil of seeing her past relationships unfold before her, decides to join their party, using her vast experience to guide them. She brutally dispatches a bandit group, showcasing her ruthless combat skills, but also experiences physical pain for the first time in decades, a sensation she finds both jarring and strangely welcome.
Upon reaching the Royal Capital, Meme strategically maneuvers to be accepted into Oscar's party during an audience with the King, a figure she despises for his past betrayals. Oscar, wanting to protect Meme, resists her involvement, leading to a tense confrontation. Meme, frustrated by Oscar's immaturity and naivety, challenges him to a duel. Though Oscar is outmatched in skill, he cleverly uses the environment to land a decisive blow, earning a symbolic victory. Meme, though outwardly dismissive, is forced to acknowledge Oscar's potential and her own diminished power, refusing his offer of a hand due to her self-perceived "dirty" soul.
The party then begins recruiting, starting with Olivia Barnett, a talented but outcast noble magic-user whom Meme knew and loved in a past life. Meme's recruitment of Olivia is tinged with bittersweet nostalgia and guilt, as he leads her towards a fate he knows too well. From a powerful, solitary warrior burdened by endless cycles of failure, Meme has transformed into a weaker, more emotionally vulnerable guide, forced to rely on others and confront his own humanity, all while relentlessly pursuing his ultimate goal.